Postdoctoral Fellow

Currently: Animal Kingdom

Formerly: Georgia State University & MD Anderson Cancer Center

Curriculum Vitae


Publications (for complete publications, including those from other labs, see below)


I worked as a post-doctoral associate with Dr. Sarah Brosnan from October 2010 until September 2011. During this time I studied responses to inequity in squirrel and owl monkeys as part of an NSF grant. In addition, I looked at the relationship between personality and responses to inequity in chimpanzees.

I received my Ph.D. in personality and social psychology from the University of Texas at Austin in 2010 working with Sam Gosling, Ph.D. My dissertation work focused on the development and validation of a chimpanzee personality rating scale using a variety of approaches and methods including behavior, endocrinology, physiology and injury and illness data. During graduate school I also worked on several personality projects with different species including humans and cheetahs that had health and well-being implications.

Following my time at GSU, I was a post-doctoral researcher position at the Lester E. Fisher Center for the Study and Conservation of Apes at the Lincoln Park Zoo. I was the project coordinator for Project ChimpCARE ( in connection with and developed by Dr. Steve Ross. The main goal of the project was to improve the well-being of chimpanzees living in captivity in the United States, particularly those living as pets, performers and in other private housing. One main line of research was a multi-institutional study of ex-pet/ex-performer chimpanzees living in zoos and sanctuaries throughout the United States.  I am currently a postdoc at Disney's Animal Kingdom.


Freeman, H.D. (in press) Using the Science of Animal Personality as a Tool for Optimizing Animal Welfare in Zoos. Connect Magazine for Association of Zoos and Aquariums.

Orban, D., Freeman, H., Wheaton, C., Mellen, J., Soltis, J., & Leighty, K. (2015). Use of Science to Enhance Animal Welfare at Disney’s Animal Kingdom. World Association of Zoos and Aquariums Magazine. 16: 18-21.

Latzman, R. D., Hecht, L.K., Freeman, H.D., Schapiro, S. J., & Hopkins, W.D. (in press). Investigating the Neuroanatomical Correlates of Personality in Chimpanzees (Pan troglodytes): Associations between Personality and Frontal Cortex. NeuroImage.

Brosnan, S.F., Hopper, L.M., Richey, S., Freeman, H.D., Talbot, C.F., Gosling, S.D., Lambeth, S.P., & Schapiro, S.J. (2015). Personality influences responses to inequity and contract in chimpanzees. Animal Behaviour. 101: 75 – 87.

Latzman, R.D., Freeman, H.D., Schapiro, S.J., & Hopkins, W.D. (2015) The contribution of genetics and early rearing experiences to hierarchical personality dimensions in chimpanees (Pan troglodytes). Journal of Personality and Social Psychology. DOI:10.1037/pspp0000040.

Latzman, R. D., Drislane, L., Hecht, L. K.*, Brislin, S., Patrick, C. J., Lilienfeld, S. O., Freeman, H.D., J., Schapiro, S. J., & Hopkins, W. D. (2015). A chimpanzee (Pan troglodytes) model of triarchic psychopathy constructs: Development and initial validation. Clinical Psychological Science.

Freeman, H.D., Ross, S.R. (2014). The Impact of Atypical Early Histories on Pet or Performer Chimpanzees. PeerJ. 2:e579; DOI 10.7717/peerj.579.

Hopper, L.M., Price, S.A., Freeman, H.D., Lambeth, S.P., Schapiro, S.J., & Kendal, R.L. (2014) Influence of personality, age, sex, and estrous state on chimpanzee problem-solving success. Animal Cognition. DOI: 10.1007/s10071-013-0715-y

Freeman, H.D., Sullivan, J., Hopper L.M., Talbot, C., Holmes A.N., Schultz-Darken, N., Williams, L.E., Brosnan, S.F., (2013) Different responses to inequity by three primate species provides support for the importance of interdependence in the selection of social behavior.  PLoS ONE. Oct 9;8(10):e76297

Reamer, L., Haller, R., Thiele, E., Freeman, H.D, Lambeth, S.P., Schapiro, S.J. (2014). Factors affecting initial training success of blood glucose testing in captive Chimpanzees (Pan troglodytes).  Zoo Biology. DOI: 10.1002/zoo.21123.

Freeman, H., Brosnan, S.F., Hopper, L.M., Lambeth, S.P., Schapiro, S.J., Gosling, S.D. (2013). A New Approach to Developing a Comprehensive and Comparative Questionnaire for Measuring Personality in Chimpanzees. American Journal of Primatology 75(10):1042-53.

Freeman, H., Gosling, S, Schapiro, S.J. (2011). Methods for assessing personality in non-human primates. In A. Weiss, J.  King, & L. Murray (Eds.), Personality and Behavioral Syndromes in Nonhuman Primates. New York: Springer.

Talbot, C.F., Freeman, H., Williams, L.E., Brosnan, S. F. (2011). Squirrel monkeys’ response to inequitable outcomes indicates a behavioural convergence within the primates. Biology Letters. 7: 680-682.

Freeman, H. Beer, J. (2010) Frontal lobe activation mediates the relation between sensation seeking and cortisol increases. Journal of Personality. 78: 497-1528.

Freeman, H., Gosling, S. (2010) Personality in non-human primates: A review and evaluation of past research. American Journal of Primatology. 71: 1-19.

Losin, E.A., Russell, J.L., Freeman, H., Meguerditchian, A., & Hopkins, W.D. (2008). Left hemisphere specialization for oro-facial movements of learned vocal signals by captive chimpanzees. PLoS ONE. 3: 1-7.

Cantalupo, C., Freeman, H., Rodes, W., Hopkins, W.D (2008). Handedness for Tool Use Correlates with Cerebellar Asymmetries in Chimpanzees (Pan troglodytes). Behavioral Neuroscience.122: 191-198.

Hopkins, W.D., Russell, J., Remkus, M., Freeman, H., & Schapiro, S.J. (2007). Handedness and grooming in chimpanzees (Pan troglodytes): Comparative analysis between findings in captive and wild individuals. International Journal of Primatology, 28, 1315-1326.

Hostetter, A.B. Russell, J. Freeman, H., Hopkins, W.D. (2006). Now you see me, now you don’t: evidence that chimpanzees understand the role of eyes in attention. Animal Cognition.10, 55-62.

Hopkins, W. D., Russell, J., Freeman, H., Reynolds, E. Griffis, C., &  Leavens, D. L. (2006). Lateralized scratching in chimpanzees (Pantroglodytes).Evidence of a Functional Asymmetry During Arousal. Emotion. 6, 553-559.

Hopkins, W.D., Cantalupo, C., Freeman, H., Russell, J., Kachin, M., Nelson, E. (2005). Chimpanzees Are Right Handed When Recording Bouts Rather Than Frequencies in Hand Use. Laterality. 10, 121-130.

Hopkins, W.D., Russell, J., Cantalupo, C., Freeman, H., Shapiro, S.J. (2005). Factors influencing throwing in chimpanzees (Pan troglodytes). Journal of Comparative Psychology. 119, 363-370.

Hopkins, W.D., Russell, J., Freeman, H., Buehler, N., Reynolds, E., Shapiro, S.J. (2005). The distribution and development of handedness for manual gestures in captive chimpanzees (Pan troglodytes). Psychological Science, 16, 487-493.

Freeman, H., Cantalupo, C., Hopkins, W.D.  (2004). Asymmetries in the hippocampus and amygdala of chimpanzees (Pan troglodytes). Behavioral Neuroscience. 118: 1460-1465.